how much signatures does comodo have and when will comodo 5 be released
Comodo has many signatures. Plus, a list of 1000000 Whitelisted files and Blacklisted files.
Comodo 5.0 is only in Beta right now - It will be released when every bug is fixed. And Security Bugs of course.
but how much sign
4,509,903 sigs. as of yesterday.
The number of signatures used by an antivirus or antimalware program is not that critical. The detection rate the the most important issue.
For a more detailed explanation of signatures, see this post.
A report from AV comparatives should be forthcoming, and this will provide a more detailed comparison with other anti-malware programs (including detection rate, false positives, etc.).
But, you should also be aware that Comodo Internet Security offers some of the strongest prevention, as shown by the tests at Matousec.